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Initially, the plan was for the voyage to be made in two vessels, the other being the smaller Speedwell. The first voyage of the ships departed Southampton, England, on August 5, 1620; but the Speedwell developed a leak, and had to be refitted at Dartmouth.
On the second attempt, the ships reached the Atlantic Ocean but again were forced to return to Dartmouth because of the Speedwell's leak.
It would later be revealed that there was in fact nothing wrong with the Speedwell. The crew had sabotaged it in order to escape the year long commitment of their contract.
After reorganisation, the final sixty-six day voyage was made by the Mayflower alone, leaving Plymouth, England on September 6. With 102 passengers plus crew, each family was allotted a very confined amount of space for personal belongings.
The ship probably had a crew of twenty-five to thirty, along with other hired personnel; however, only the names of five are known, including John Alden. William Bradford, who penned our only account of the Mayflower voyage, wrote that John Alden (archaic spellings) "was hired for a cooper [barrel-maker], at South-Hampton, where the ship victuled; and being a hopefull yong man, was much desired, but left to his owne liking to go or stay when he came here; but he stayed, and maryed here."
The intended destination was an area near the Hudson River, in "North Virginia". However the ship was forced far off-course by inclement weather and drifted well north of the intended Virginia settlement. As a result of the delay, the settlers did not arrive in Cape Cod till the onset of a harsh New England winter. The settlers ultimately failed to reach Virginia where they had already obtained permission from the London Company to settle. To establish legal order and to quell increasing strife within the ranks, the settlers wrote and signed the Mayflower Compact after the ship dropped anchor at the tip of Cape Cod on November 11, in what is now Provincetown Harbor.
The settlers, upon initially setting anchor, explored the snow-covered area and discovered an empty Native American village. The curious settlers dug up some artificially-made mounds, some of which had stored corn while others were burial sites. The settlers stole the corn, sparking friction with the locals. They explored the area of Cape Cod for several weeks and decided to relocate after a difficult encounter with the local native Americans, the Nausets.
During the winter the passengers remained on board the 'Mayflower', suffering an outbreak of a contagious disease described as a mixture of scurvy, pneumonia and tuberculosis. When it ended, there were only 53 persons still alive, half of the passengers and half of the crew. In spring, they built huts ashore, and on March 21, 1621, the surviving passengers left the 'Mayflower'.
On April 5, 1621, the Mayflower set sail from Plymouth to return to England, where she arrived on May 6, 1621.
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